I'm still saying that they were at some point overlaid with color, post production...if you look at the left side of the scan just at the pictures border, you can actually see the black and white base card where the faint green tint missed the edge, also, you can see where that same green addition actually dripped a little at the bottom right corner...
I'm no expert in printing technique or technology, and I agree there's almost no way someone did that with brushes by hand, but it seems pretty obvious that this series is clearly a post production creation and not an original color copy created by Gum Inc
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